Douglas J. Futuyma is Distinguished Professor in the Department of Ecology and Evolution at the State University of New York at Stony Brook. He received his B.S. from Cornell University and his Ph. D. in Zoology at the University of Michigan with Lawrence Slobodkin. Dr. Futuyma is the author of three previous editions of Evolution, as well as three editions of its predecessor, Evolutionary Biology. He received the 1997 Sewall Wright Award of the American Society of Naturalists and the 2012 Joseph Leidy Award of the Academy of Natural Sciences of Drexel University (Philadelphia). Dr. Futuyma has served as President of the Society for the Study of Evolution, the American Society of Naturalists, and the American Institute of Biological Sciences, and was elected a Fellow of the American Academy of Arts and Sciences in 1996 and the National Academy of Sciences in 2006. Mark Kirkpatrick is the Painter Centennial Professor of Genetics in the Department of Integrative Biology at the University of Texas at Austin. He received his B.A. in Biology from Harvard in 1978 and his Ph.D. in Zoology from the University of Washington with Monty Slatkin in 1983. Dr. Kirkpatrick has received a Guggenheim Fellowship (1997) and a Poste Rouge Fellowship (France, 1997). He is a Fellow of the American Academy of Arts and Sciences (2008), and of the American Association for the Advancement of Science (2016). Dr. Kirkpatrick received the Sewall Wright Award from the American Society of Naturalists (2014). He has served as Associate Editor of The American Naturalist, Theoretical Population Biology, and Genetics, and on the Editorial Boards of The Annual Review of Ecology, Evolution, and Systematics and Proceedings of the Royal Society of London. Dr. Kirkpatrick's research interests are in evolutionary genetics.